Megasporogenesis,Microsporogenesis and Development of Female and Male Gametophyte of Tricyrtis maculata(D.Don)Machride

The megasporogenesis,microsporogenesis and development of female and male gametophyte of Tricyrtis maculata(D.Don)Machride were observed by the method of paraffin sections.The main results as follows:(1)The anther is four sporangiate and the development of anther wall is the monocotyledonous type.The anther wall is composed of epidermis,endothecium,middle layer and tapetum.The epidermise consist of one layer of cells and still existed until the anther matured.The endothecium developed bandingshaped thickenings when the anther matured.The young anther wall has 2~3layer middle layers and degenerated when maturity of the anther.The tapetum belongs to secretory type and degenerated at the time of anther matured.(2)The cytokinesis of the microspore mother cells in meiosis is successive type.The microspore tetrads are tetrahedral and isobilateral spore.The mature pollen grain is the 2-celled type.(3)The ovary has three chamber which has more ovule that is double integuments,tenuinucellate and anatropous.The archesporium directly functions as the megaspore mother cell.After meiosis,the megasporocyte divides into linear or T-shaped type tetrads.The megaspore in chalaza end usually is functional one.The embryo sac is a polygonum type.(4)There are some abnormal phenomena in the development of the anther that bring about the pollen degeneration.