Biology of Amphibians

Introduction to amphibia - the world of amphibians, historical resume, prospects for the future. Part 1 Life History: reproductive strategies - reproductive cycles, reproductive mode, quantitative aspects, parental care, evolution of reproductive strategies courtship and mating - location of breeding site, secondary sexual characters, courtship behaviour, fertilization and oviposition, sexual selection, evolution of mating systems vocalization - anuran communication system, mechanisms of sound production and reception, kinds of vocalizations and their functions, abiotic factors affecting vocalization, interspecific significance of vocalization, phylogenetic implications of vocalization eggs and development - spermatozoa and fertilization, egg structure, egg development, hatching and birth, development and amphibian diversity larvae - morphology of larvae, adaptive types of larvae, physiology and ecology, social behaviour, evolutionary significance of larvae metamorphosis - endocrine control, other biochemical changes, morphological changes, neoteny, ecological and evolutionary significance of metamorphosis. Part 2 Ecology: relationships with the environment - water economy, temperature, gas exchange, energy metabolism and energy budgets, ecological synthesis food and feeding - prey selection, location of prey, capture of prey, evolution of prey-capturing mechanisms and strategies enemies and defence - diseases, parasites, predators, anti-predator mechanisms, evolution of defence mechanisms population biology - characteristics of individuals, movements and territoriality, demography, factors regulating populations community ecology and species diversity - community structure, species diversity, evolution of amphibian communities. Part 3 Morphology: musculoskeletal system - skull and hyobranchium, axial system, appendicular system, integration of functional units integumentary, sensory and visceral systems - integument, sensory receptor systems, nervous system, circulatory and respiratory systems, urogenital system, digestive system, endocrine glands, evolutionary considerations. Part 4 Evolution: origin and early evolution - nature of a tetrapod, primitive tetrapods, tetrapod affinities (lungfishes or lobe-fins?), diversity and evolution of early tetrapods, status of the lissamphibia cytogenetic, molecular and genomic evolution - cytogenetics, molecular evolution, genomic evolution phylogeny - caudata, gymnophiona, anura biogeography - biogeographic principles, historical setting, lissamphibia, caudata, gymnophiona, anura classification.