Vegetation of the Stipa loess steppe in Ningxia (northern China) in relation to grazing intensity

Abstract Large areas of northern and western China are covered with steppe vegetation which is grazed with different intensities. Areas which are not or only slightly grazed are rare and mostly exist in nature reserves. We investigated floristic composition and species diversity of the steppe vegetation of the autonomous region Ningxia Hui with emphasis on the Yunwushan dry steppe nature reserve in relation to grazing intensity; all 77 plots were analyzed using ordination techniques, and the relationship between the variation in species composition and environmental conditions was analyzed. In Ningxia, two types of loess grass steppes prevail: in areas with lower altitudes from approximately 1650–1950 m a.s.l., the Stipa bungeana steppe occurs whereas the Stipa grandis steppe is typical for higher altitudes between 1900 and 2100 m a.s.l. For both vegetation types, three subtypes can be distinguished according to different levels of grazing intensity. With increasing grazing intensity, the number of plant species per plot strongly declines and the typical steppe vegetation types are dominated by Artemisia species (A. frigida in the Stipa grandis community and A. sacrorum in the Stipa bungeana community). In contrast to studies carried out in neighboring areas, in our study area the Artemisia steppes do not represent the final state of degradation. Instead, Convolvulus ammanii dominates in heavily grazed areas of both Stipa steppe types. Detrended correspondence analysis confirms the classification and shows that the heavily grazed subtypes of the Stipa grandis and the Stipa bungeana steppe with Convolvulus ammanii are more similar to each other than the other vegetation types. The first axis of the detrended correspondence analysis is strongly related to the grazing intensity (r = 0.93). Our results show that grazing intensity is an important factor for the differentiation of the steppe grasslands and a severe threat to species diversity.